unfort - as you've seen the 'cheaper' lenses tend to be dark,. around 700 quid is the sigma 50-500mm F4-6.3 EX DG (86mm) it's meant to be pretty good optically perhaps the sigma APO 170-500mm F5-6.3 DG is similar / better even
once you go past 200 or 300 mm you are either needing something with image stabilisation (for hand held) or expect to use a tripod to get a sharp image,. esp if you are forced to use slower shutter speeds because of the f stop
prime lenses will give you more quality but you can loose a lot of that once you put a tele converter on it,.
something like the nikon 80-400 has VR - it's good optically (from reports I don't have one) and it has VR (nikons image stabalisation system) it's not cheap though..
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I have checked it out and if I had to choose between the two I would pick the faster one i.e 135-400 f4.5-5.6. I gives you a little more in terms of light compared to the 170-500.
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I don't know but you might find the 170-500 is 5.6 at 400mm,. I would go for the 500mm on spec esp if 500 is going to be long enough and you were thinking of using a tele converter on the 400mm to get to 500mm
Sil